From the article:

The researchers found an average of around 100 microplastic particles per liter in glass bottles of soft drinks, lemonade, iced tea and beer. That was five to 50 times higher than the rate detected in plastic bottles or metal cans.

...

"We then noticed that in the glass, the particles emerging from the samples were the same shape, color and polymer composition—so therefore the same plastic—as the paint on the outside of the caps that seal the glass bottles," she said.

This could then "release particles onto the surface of the caps," it added.